Why does my webservice application on WAS 8.5.5.10 only run on one cluster member, but fails on another?

We have a WAS ND cell 8.5.5.10 with two cluster members.
There is a webservices application deployed to the cluster.

But when we try to access the wsdl of the webservice (e.g. with the full URL from in a browser), then we can only access the wsdl on the first node, but it fails on the second one (HTTP Status Code 404).

Node Synchronization is fine, the application appears to be started on the second node as well.

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